I would say roughly 30-40% in Substance Modeler and 60-70% including all the keying and final prep for print in Zbrush.
It was a back and forth. The armour, for instance, was blocked out in ZB > larger detailing and further shape refinements in Modeler > back to ZB for final polishing/surface detailing and print prep. Most of the larger detailing from Modeler got an extra surfacing pass in Zbrush.
Another example is the hair: initial blockout in Modeler > final clump and strand adjustments and strand sculpting and refinements in ZB.
So yeah, it was a back and forth using the strength of each program. Once the ZB scene was setup to scale it was easy to jump in and out between the 2 programs.